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Eumaeina Doubleday 1847

Andrew V. Z. Brower
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taxon links [up-->]Symbiopsis [up-->]Magnastigma [up-->]Cyanophrys [up-->]Theclopsis [up-->]Terenthina [up-->]Thestius [up-->]Janthecla [up-->]Paiwarria [up-->]Erora [up-->]Trichonis [up-->]Evenus [up-->]Thereus [up-->]Electrostrymon [up-->]Laothus [up-->]Balintus [up-->]Kolana [up-->]Podanotum [up-->]Lathecla [up-->]Lamasina [up-->]Enos [up-->]Aubergina [up-->]Ostrinotes [up-->]Siderus [up-->]Hypostrymon [up-->]Apuecla [up-->]Phaeostrymon [up-->]Temecla [up-->]Strephonota [up-->]Marachina [up-->]Exorbaetta [up-->]Camissecla [up-->]Arcas [up-->]Chalybs [up-->]Satyrium [up-->]Atlides [up-->]Ignata [up-->]Bistonina [up-->]Nesiostrymon [up-->]Arawacus [up-->]Ocaria [up-->]Micandra [up-->]Olynthus [up-->]Theritas [up-->]Contrafacia [up-->]Calycopis [up-->]Johnsonita [up-->]Thepytus [up-->]Megathecla [up-->]Timaeta [up-->]Allosmaitia [up-->]Ziegleria [up-->]Arumecla [up-->]Chlorostrymon [up-->]Semonina [up-->]Callophrys [up-->]Rekoa [up-->]Dicya [up-->]Phothecla [up-->]Panthiades [up-->]Oenomaus [up-->]Busbiina [up-->]Brevianta [up-->]Thaeides [up-->]Eumaeus [up-->]Brangas [up-->]Mithras [up-->]Iaspis [up-->]Gargina [up-->]Salazaria [up-->]Nicolaea [up-->]Lamprospilus [up-->]Porthecla [up-->]Strymon [up-->]Tmolus [up-->]Rhamma [up-->]Parrhasius [up-->]Ipidecla [up-->]Celmia [up-->]Pseudolycaena [up-->]Ministrymon [up-->]Michaelus [up-->]Theorema [up-->]Penaincisalia [down<--]Theclini Interpreting the tree
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The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

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Containing group: Theclini


A diverse subtribe, with its greatest diversity (over 1000 species) in the neotropics.

Discussion of Phylogenetic Relationships

Sections of Eumaeina are based on the classification of Robbins and Lamas (2004). Many of the species, particularly the neotropical ones, were originally described in the catch-all genus Thecla, which is now restricted to a group of mainly Palearctic species in a separate subtribe.


Bridges, C. A. 1988 Catalogue of family-group and genus-group names (Lepidoptera: Rhopalocera). Urbana, IL: Charles A. Bridges.

Eliot, J. N. 1973 The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement. Bull. Br. Mus. Nat. Hist. (Ent.) 28, 371-505, 6 pls.

Robbins, R. K. and G. Lamas. 2004 Lycaenidae. in Lamas, G. (ed.) Atlas of Neotropical Lepidoptera. Checklist: Part 4A Hesperioidea - Papiionoidea. Gainesville: Scientific Publishers/Association of Tropical Lepidoptera.

Title Illustrations
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Scientific Name Callophrys xami
Location Mexico City, Mexico
Comments Laying eggs on succulent (Larval foodplant)
Specimen Condition Live Specimen
Sex Female
Life Cycle Stage Adult
Source Collection Maryposa
Copyright © Maria Franco
Scientific Name Mithras nautes
Location Brazil: Rond?nia, Poto Velho, Rancho Grande
Specimen Condition Live Specimen
Identified By Hank Brodkin
Life Cycle Stage adult
View ventral
Source Nautes Hairstreak (Mithras nautes)
Source Collection Neotropical Butterflies
Copyright © 2006 Hank Brodkin
Scientific Name Eumaeus atala
Location Palm Beach County, Florida, USA
Specimen Condition Live Specimen
Identified By Patrick Coin
Behavior Female ovipositing on hostplant, (Zamia pumila, a cycad).
Sex Female
Life Cycle Stage adult (imago)
View lateral
Source Photo#5432
Source Collection BugGuide.Net
ToL Image Use creative commons This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 2.5.
Copyright © 2004 Patrick Coin
About This Page

Middle Tennessee State University, Murfreesboro, Tennessee, USA

Correspondence regarding this page should be directed to Andrew V. Z. Brower at

Page: Tree of Life Eumaeina Doubleday 1847. Authored by Andrew V. Z. Brower. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Brower, Andrew V. Z. 2008. Eumaeina Doubleday 1847. Version 24 March 2008 (under construction). http://tolweb.org/Eumaeina/111502/2008.03.24 in The Tree of Life Web Project, http://tolweb.org/

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