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This is an archived version of a Tree of Life page. For up-to-date information, please refer to the current version of this page.


taxon links [down<--]Carabidae_Conjunctae [up-->]Pterostichini [up-->]Pseudomorphini Not Monophyletic [up-->]Morionini [up-->]Cnemalobini [up-->]Catapieseini [up-->]Platynini [up-->]Zabrini [up-->]Bascanini [up-->]Peleciini [up-->]Cuneipectini [up-->]Licinini [up-->]Oodini [up-->]Panagaeini [up-->]Chlaeniini [up-->]Harpalini [up-->]Dryptini [up-->]Galeritini [up-->]Helluonini [up-->]Idiomorphini [up-->]Orthogoniini [up-->]Amorphomerini [up-->]Lebiomorpha Interpreting the tree
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This tree diagram shows the relationships between several groups of organisms.

The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

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You can click on the root to travel down the Tree of Life all the way to the root of all Life, and you can click on the names of descendent subgroups to travel up the Tree of Life all the way to individual species.

For more information on ToL tree formatting, please see Interpreting the Tree or Classification. To learn more about phylogenetic trees, please visit our Phylogenetic Biology pages.

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Containing group: Carabidae Conjunctae


The Harpalinae comprise those carabids with conjunct mesocoxae (as explained on the page for the Carabidae Conjunctae, and which also have the following features: In addition, members of this group have the following properties:
Title Illustrations

From left to right,

Photographs copyright © 1995, David R. Maddison

Scientific Name Pterostichus sp.
Location Florida
Copyright © 1995 David R. Maddison
Scientific Name Chlaenius pimalicus
Location Arizona
Copyright © 1995 David R. Maddison
Scientific Name Galerita atripes
Location Arizona
Copyright © 1995 David R. Maddison
Scientific Name Lebia bivittata
Location Arizona, U.S.A.
Copyright © 1995 David R. Maddison
About This Page

Correspondence regarding this page should be directed to David R. Maddison at

Citing this page:

Maddison, David R. 1995. Harpalinae. Version 01 January 1995 (under construction). http://tolweb.org/Harpalinae/100/1995.01.01 in The Tree of Life Web Project, http://tolweb.org/

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This page is a Tree of Life Branch Page.

Each ToL branch page provides a synopsis of the characteristics of a group of organisms representing a branch of the Tree of Life. The major distinction between a branch and a leaf of the Tree of Life is that each branch can be further subdivided into descendent branches, that is, subgroups representing distinct genetic lineages.

For a more detailed explanation of the different ToL page types, have a look at the Structure of the Tree of Life page.

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