Whitefoot Fern, White-foot LipfernCarl Rothfels
Cheilanthes leucopoda is a small- to medium-sized fern with pentagonal glandular-hairy leaves. It occurs in central and northern Mexico, where it can be common, and penetrates into the United States in the Edwards Plateau of Texas (where it is rare—S2; NatureServe, 2008). It shares many features with the species of the myriopterid clade (leaf margins differentiated and recurved into false indusia, discontinuous sori concentrated on the segment tips, etc.), but is only distantly related to them, and will require a new name (either under Notholaena, or as a new genus; Rothfels et al., 2008).
Cheilanthes leucopoda is a species of limestone cliffs and slopes, at middle elevations. Its distinct 4-pinnate pentagonal leaves are borne on thick but weak, straw-colored stalks that are often twice as long as the leaf blades themselves, giving the plants a rather umbrella-like appearance. The leaves and stalks are densely covered with short glandular hairs.
Gastony, G. J., and D. R. Rollo. 1995. Phylogeny and generic circumscriptions of cheilanthoid ferns (Pteridaceae: Cheilanthoideae) inferred from rbcL nucleotide sequences. American Fern Journal 85:341-360.
Gastony, G. J., and D. R. Rollo. 1998. Cheilanthoid ferns (Pteridaceae: Cheilanthoideae) in the southwestern United States and adjacent Mexico--A molecular phylogenetic reassessment of generic lines. Aliso 17:131-144.
Kramer, K. U. 1990. Notes on the higher level classification of the recent ferns. Pages 49--52 in The Families and Genera of Vascular Plants (K. U. Kramer, and P. S. Green, eds.). Springer-Verlag, Berlin.
Mickel, J. T. 1979. The fern genus Cheilanthes in continental United States. Phytologia 41:431-437.
Mickel, J. T., and A. R. Smith. 2004. The Pteridophytes of Mexico. The New York Botanical Garden Press, New York.
NatureServe. 2008. NatureServe Explorer, Arlington, Virginia.
Rothfels, C. J., M. D. Windham, A. L. Grusz, G. J. Gastony, and K. M. Pryer. 2008. Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns Taxon 57:712-724.
Schuettpelz, E., H. Schneider, L. Huiet, M. D. Windham, and
Tryon, R. M., and A. F. Tryon. 1973. Geography, spores, and evolutionary relations in the cheilanthoid ferns. Pages 145-153 in The phylogeny and classification of ferns. (A. C. Jermy, J. A. Crabbe, and B. A. Thomas, eds.). Academic Press, New York.
Tryon, R. M., and A. F. Tryon. 1982. Ferns and Allied Plants with Special Reference to Tropical America. Springer-Verlag, New York.
Tryon, R. M., A. F. Tryon, and K. U. Kramer. 1990. Pteridaceae. Pages 404 in The Families and Genera of Vascular Plants (K. U. Kramer, and P. S. Green, eds.). Springer-Verlag, Berlin.
Windham, M. D. 1993b. Pteridaceae. Pages 122--124 in Flora of North America (Flora of North American Editorial Committee, ed.) Oxford University Press, New York.
Windham, M. D., L. Huiet, E. Schuettpelz, C. J. Rothfels, J. Beck, A. L. Grusz, G. Yatskievych, and K. M. Pryer. 2008. Using plastid and nuclear DNA sequences to redraw generic boundaries and demystify species complexes in cheilanthoid ferns. American Fern Journal (in review).
Windham, M. D., and G. Yatskievych. 2003. Chromosome studies of cheilanthoid ferns (Pteridaceae: Cheilanthoideae) from the western United States and Mexico. American Journal of Botany 90:1788-1800.
Duke University, Durham, North Carolina, USA
Correspondence regarding this page should be directed to Carl Rothfels at
Page copyright © 2008 Carl Rothfels
Page: Tree of Life Cheilanthes leucopoda Authored by . Whitefoot Fern, White-foot Lipfern.Carl Rothfels. The TEXT of this page is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.
- First online 23 December 2008
- Content changed 23 December 2008
Citing this page:
Rothfels, Carl. 2008. Cheilanthes leucopoda http://tolweb.org/Cheilanthes_leucopoda/133578/2008.12.23 in The Tree of Life Web Project, http://tolweb.org/. Whitefoot Fern, White-foot Lipfern. Version 23 December 2008 (under construction).