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Harpalinae

Kipling Will and David R. Maddison
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taxon links [up-->]Helluonini [up-->]Amorphomerini [up-->]Peleciini [up-->]Idiomorphini [up-->]Harpalini [up-->]Physocrotaphini [up-->]Oodini [up-->]pterostichite grade [up-->]Zuphiini [up-->]Pseudomorphini [up-->]Geobaenini [up-->]Anthiini [up-->]Orthogoniini [up-->]Chlaeniini [up-->]Bascanini [up-->]Panagaeini [up-->]Dercylini [up-->]Licinini [up-->]Lebiomorpha [up-->]Chaetogenyini [up-->]Hexagoniini [up-->]Dryptini [up-->]Galeritini [up-->]Ctenodactylini [up-->]Platynini [up-->]Ginema thomasi Not MonophyleticMonophyly Uncertain[down<--]Carabidae Conjunctae Interpreting the tree
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The root of the current tree connects the organisms featured in this tree to their containing group and the rest of the Tree of Life. The basal branching point in the tree represents the ancestor of the other groups in the tree. This ancestor diversified over time into several descendent subgroups, which are represented as internal nodes and terminal taxa to the right.

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This tree is a composit of various published and unpublished data.
Containing group: Carabidae Conjunctae

Introduction

This large clade contains more than half of carabid species.  It is supported as monophyletic by both morphological and molecular data (Maddison et al., 1999; Ober, 2002).

Characteristics

The Harpalinae comprise those carabids with conjunct mesocoxae (as explained on the page for the Carabidae Conjunctae, and which also have the following features: In addition, members of this group have the following properties:

References

Maddison, D.R., M.D. Baker, and K.A. Ober. 1999. Phylogeny of carabid beetles as inferred from 18S ribosomal DNA (Coleoptera: Carabidae). Systematic Entomology, 24:103-138.

Ober, K.A. 2002. Phylogenetic relationships of the carabid subfamily Harpalinae (Coleoptera) based on molecular sequence data. Molecular Phylogenetics and Evolution, 24: 228-248.

Title Illustrations
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Scientific Name Pterostichus morionides
Identified By K.Will
Life Cycle Stage Adult
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © Kipling Will
Scientific Name Calybe sallei
Location USA: Texas: Fredonia
Specimen Condition Dead Specimen
Sex female
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 2005 David R. Maddison
Scientific Name Bradycellus
Location USA: New Mexico: Grant Co., Gila River, Billings Vista
Specimen Condition Dead Specimen
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 2005 David R. Maddison
Scientific Name Chlaenius ruficauda
Location USA: New Mexico: Grant Co., Gila River, Billings Vista
Specimen Condition Dead Specimen
Identified By David Maddison
Sex Male
Image Use creative commons This media file is licensed under the Creative Commons Attribution License - Version 3.0.
Copyright © 2002 David R. Maddison
About This Page

Kipling Will
University of California, Berkeley, California, USA

David R. Maddison
Oregon State University

Correspondence regarding this page should be directed to Kipling Will at and David R. Maddison at

Page: Tree of Life Harpalinae. Authored by Kipling Will and David R. Maddison. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Will, Kipling and David R. Maddison. 2006. Harpalinae. Version 07 July 2006 (under construction). http://tolweb.org/Harpalinae/100/2006.07.07 in The Tree of Life Web Project, http://tolweb.org/

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Harpalinae

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