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Protococcidiorida Kheisin 1956

Jan Šlapeta
Containing group: Apicomplexa


This group of apicomplexan parasites is characterised by the presence of extracellular gamogony and sporogony. Merogony is apparently absent. Extracellular sporozoites develop directly into gamonts. These parasites are found in estuarine, marine and tidal annelids. In taxonomical texts they are classified as Order Protococcidiida Kheisin, 1956.

There are four families:

Family: Eleutheroschizonidae Chaton & Villeneuve, 1936
Family: Myriosporidae Grasse, 1953
Family: Angeiocystidae Leger, 1911
Family: Grellidae Levine, 1973

Grellia dinophili sporozoites in the body cavity of the marine archiannelid Dinophilus gyrociliatus were studied using transmission electron microscopy.  They possess a distinct crystalloid body and a nice set of rhoptries (Elwasila 1983).

Nothing is known about the genetic identity of these organisms.

Other Names for Protococcidiorida Kheisin 1956


Elwasila, M. 1983. A fine-Structural comparison of the Sporozoites of Grellia (Eucoccidium) dinophili in Dinophilus gyrociliatus and of Isospora fells. in the Mouse. Z Parasitenkd 69:583-589

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This page is being developed as part of the Tree of Life Web Project Protist Diversity Workshop, co-sponsored by the Canadian Institute for Advanced Research (CIFAR) program in Integrated Microbial Biodiversity and the Tula Foundation.

Jan Šlapeta
Faculty of Veterinary Science, University of Sydney, New South Wales, Australia

Correspondence regarding this page should be directed to Jan Šlapeta at

Page: Tree of Life Protococcidiorida Kheisin 1956. Authored by Jan Šlapeta. The TEXT of this page is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Šlapeta, Jan. 2011. Protococcidiorida Kheisin 1956. Version 18 May 2011 (under construction). http://tolweb.org/Protococcidiorida/124804/2011.05.18 in The Tree of Life Web Project, http://tolweb.org/

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