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Bembidion Latreille 1802

David R. Maddison
Containing group: Bembidiina


This is a large genus of over 1000 described species. Most species are found in north temperate regions, although two smaller centers of diversity exist in the southern hemisphere (the Andes mountains of South America, as well as New Zealand).

Adults are common, small predators and scavengers along bodies of water, from rapidly flowing mountain streams to lowland ponds. Several species living in more xeric conditions, including deserts.


Some pictures of Bembidion structure are shown here.


Erwin, T. L.  1982. Small terrestrial ground-beetles of Central America (Carabidae: Bembidiina and Anillina). Proceedings of the California Academy of Sciences, 42: 455–496.

Erwin, T. L.  1984. Studies of the Tribe Bembidiini (Coleoptera: Carabidae): Lectotype designations and species group assignments for Bembidion species described by Thomas L. Casey and others. Pan-Pacific Entomologist, 60: 165–187.

Erwin, T. L., and D. H. Kavanaugh. 1981. Systematics and zoogeography of Bembidion Latreille: I. The carlhi and erasum groups of western North America (Coleoptera: Carabidae, Bembidiini). Entomologica Scandinavica Supplement, 15: 33–72.

Lindroth, C.H.  1963. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 3. Opuscula Entomologica, Supplementum XXIV, pp. 201–408.

Lindroth, C.H.  1976. Genus Bembidion Latreille (Coleoptera: Carabidae) in New Zealand: a revision. New Zealand Journal of Zoology, 3: 161–198.

Lindroth, C.H.  1980. A revisionary study of the taxon Cillenus Samouelle, 1819 and related forms (Coeloptera [sic]: Carabidae, Bembidiini). Entomologica Scandinavica, 11: 179–205.

Maddison, D. R. 1985. Chromosomal diversity and evolution in the ground beetle genus Bembidion and related taxa (Coleoptera: Carabidae: Trechitae). Genetica 66(2): 93-114.

Maddison, D. R. 1993. Systematics of the Holarctic beetle subgenus Bracteon and related Bembidion (Coleoptera: Carabidae). Bulletin of the Museum of Comparative Zoology 153(3): 143-299.

Müller, J. 1918. Bestimmungstabelle der Bembidion-Arten Europas und des Mittelmeergebietes. Koleopterologische Rundschau, 7: 26–117.

Müller, J.  1926–1927. Tableaux de détermination des coléoptères d'Europe. Heft LXXXIX (89). Carabidae. Genre Bembidium Latr. d'Europe et de la région méditerranéenne. Miscellanea Entomologica, 29–30: 1–100. [French translation of Müller (1918).]

Netolitzky, F.  1942, 1943. Bestimmungstabelle der Bembidion-Arten des paläarktischen Gebietes. Koleopterologische Rundschau, 28: 29–124, 29: 1–70.

Perrault, G. G. 1981. Etudes sur la tribu des Bembidiini (Coleoptera, Carabidae) I. Notes sur la classification supraspécifique. Nouvelle Revue d'Entomologie, 11: 237–250.

Perrault, G. G.  1982. Etudes sur la tribu des Bembidiini s. str. (Coleoptera, Carabidae). II. Révision du sous-genre Cyclolopha (Casey). Entomologica Basiliensia, 7: 89–126.

Schuler, L. 1959. Le genre Microserrulula Netolitzky (Bembidiini Jeannel, Col. Trechidae) les formes multispires. Revue française d'Entomologie, 26: 96–105.

Serrano, J. 1981a. A chromosome study of Spanish Bembidiidae and other Caraboidea. Genetica 57: 119-129.

Information on the Internet

Title Illustrations
Scientific Name Bembidion levettei
Location Canada: British Columbia: Malakwa
Sex Male
Copyright © 2004 David R. Maddison
Scientific Name Bembidion texanum
Location USA: Illinois: Danville
Sex Male
Copyright © 2004 David R. Maddison
Scientific Name Bembidion dorsale
Location Canada: Alberta: Oldman River at Taber
Sex Male
Copyright © 2004 David R. Maddison
About This Page

David R. Maddison
University of Arizona, Tucson, Arizona, USA

Correspondence regarding this page should be directed to David R. Maddison at

Citing this page:

Maddison, David R. 2006. Bembidion Latreille 1802. Version 17 June 2006 (under construction). http://tolweb.org/Bembidion/194/2006.06.17 in The Tree of Life Web Project, http://tolweb.org

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