Notholaena copelandii
Copeland's Cloakfern
Carl Rothfels
Introduction
Often treated as N. candida var. copelandii, Notholaena copelandii differs from N. candida in morphology (Hall, 1950), farina chemistry (Wollenweber, 1984; Wollenweber et. al 2001), geographic range, and plastid DNA sequences (Rothfels et al., 2008). I thus follow Windham (1993a) and Mickel and Smith (2004) in treating them as distinct species.
Notholaena copelandii is a species of southern Texas and northeastern Mexico, where it occurs on limestone cliffs and outcrops.
Characteristics
The morphological characters distinguishing N. copelandii from N. candida include: basal pinnae shape (a single exaggerated basioscopic pinnule on N. copelandii; pinnules gradually tapering in length in N. candida); leaf tip shape (abruptly tapered vs. gradually tapered); and rhizome scales (ciliate vs. slightly toothed). Notholaena copelandii is also similar to N. jacalenis, but the later species has much narrower leaves.
Other Names for Notholaena copelandii
- Notholaena candida var. copelandii
- Vernacular Names: Copeland's Cloakfern
References
Giauque, M. F. A. 1949. Wax glands and prothallia. American Fern Journal 39:33-35.
Hall, C. C. 1950. Notholaena copelandii, a newly recognized species of the Texano-Mexican region. American Fern Journal 40:178--187.
Mickel, J. T., and A. R. Smith. 2004. The Pteridophytes of Mexico. The New York Botanical Garden Press, New York.
NatureServe. 2008. NatureServe Explorer, Arlington, Virginia. www.natureserve.org/explorer/
Rothfels, C. J., M. D. Windham, A. L. Grusz, G. J. Gastony, and K. M. Pryer. 2008. Toward a monophyletic Notholaena (Pteridaceae): Resolving patterns of evolutionary convergence in xeric-adapted ferns Taxon 57:712-724.
Tryon, R. M. 1956. A revision of the American species of Notholaena. Contributions from the Gray Herbarium 179:1-106.
Windham, M. D. 1993a. Notholaena. Pages 143--149 in Flora of North America (Flora of North American Editorial Committee, ed.) Oxford University Press, New York.
Windham, M. D., and G. Yatskievych. 2003. Chromosome studies of cheilanthoid ferns (Pteridaceae: Cheilanthoideae) from the western United States and Mexico. American Journal of Botany 90:1788-1800.
Wollenweber, E. 1984. Exudate flavonoids of Mexican ferns as chemotaxonomic markers. Rev. Latinoamer. Quim. 15:3-11.
Wollenweber, E., M. Dörr, and J. F. Stevens. 2001. A dihydroflavonol with taxonomic significance from the fern Notholaena sulphurea. Zeitschrift für Naturforschung 56:499-502.
Title Illustrations
| Scientific Name | Notholaena copelandii |
|---|---|
| Location | USA, Texas |
| Specimen Condition | Live Specimen |
| Identified By | C.J.Rothfels |
| Image Use |
 This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
|
| Copyright |
© 2008 Carl Rothfels
|
| Scientific Name | Notholaena copelandii |
|---|---|
| Location | USA, Texas |
| Specimen Condition | Live Specimen |
| Identified By | C.J.Rothfels |
| Life Cycle Stage | sporophyte |
| Image Use |
This media file is licensed under the Creative Commons Attribution-NonCommercial License - Version 3.0.
|
| Copyright |
© 2008 Carl Rothfels
|
About This Page
Carl Rothfels
Duke University, Durham, North Carolina, USA
Correspondence regarding this page should be directed to Carl Rothfels at
Page copyright © 2008 Carl Rothfels
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- First online 23 December 2008
- Content changed 23 December 2008
Citing this page:
Rothfels, Carl. 2008. Notholaena copelandii . Copeland's Cloakfern. Version 23 December 2008 (under construction). http://tolweb.org/Notholaena_copelandii/133605/2008.12.23 in The Tree of Life Web Project, http://tolweb.org/
