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Ursus arctos

Brown Bear

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Kodiak Brown Bear
Containing group: Ursidae

Other Names for Ursus arctos


Calvignac, S, Hughes S, Tougard C, Michaux J, Thevenot M, Philippe M, Hamdine , Hanni C. 2008. Ancient DNA evidence for the loss of a highly divergent brown bear clade during historical times. Molecular Ecology 17(8):1962-1970.

Miller CR and Waits LP. 2003. The history of effective population size and genetic diversity in the Yellowstone grizzly (Ursus arctos): Implications for conservation. Proceedings of the National Academy of Sciences (USA) 100(7):4334-4339.

Shields, GF, Adams, D, Garner, G, Labelle, M, Pietsch, J, Ramsay, M, Schwartz, C, Titus, K, Williamson, S. 2000. Phylogeography of mitochondrial DNA variation in brown bears and polar bears. Molecular Phylogenetics and Evolution 15:319-326.

Taberlet P, Bouvet J. 1994. Mitochondrial DNA polymorphism, phylogeography, and conservation genetics of the brown bear Ursus arctos in Europe. Proc R Soc Lond B Biol Sci 255:195-200.

Talbot, SL, Shields, GF. 1996. Phylogeography of brown bears (Ursus arctos) of Alaska and paraphyly within the Ursidae. Molecular Phylogenetics and Evolution 5:477-494.

Valdiosera, CE, Garcia-Garitagoitia, JL, Garcia N, Doadrio I, Thomas MG, Hanni C, Arsuaga JL, Barnes I, Hofreiter M, Orlando L, Gotherstorm A. 2008. Surprising migration and population size dynamics in ancient Iberian brown bears (Ursus arctos). Proceedings of the National Academy of Sciences (USA) 105(13):5123-5128.

Waits, LP, Talbot, SL, Ward, RH, Shields, GF. 1998. Mitochondrial DNA phylogeography of the North American brown bear and implications for conservation. Conservation Biology 12(2):408-417.

Title Illustrations
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Click on an image to view larger version & data in a new window
Kodiak Brown Bear
Scientific Name Ursus arctos middendorffi
Location Kodiak National Wildlife Refuge, Alaska, USA
Comments Kodiak Brown Bear
Creator John and Karen Hollingsworth
Specimen Condition Live Specimen
Source Kodiak Brown Bear
Source Collection U.S. Fish and Wildlife Service Online Digital Media Library
Scientific Name Ursus arctos horribilis
Location Yellowstone National Park, Wyoming, USA
Specimen Condition Live Specimen
Source Grizzly Bear
Source Collection Flickr
Copyright © 2008 Jim Scarff
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Citing this page:

Tree of Life Web Project. 2008. Ursus arctos. Brown Bear. Version 12 October 2008 (temporary). http://tolweb.org/Ursus_arctos/123666/2008.10.12 in The Tree of Life Web Project, http://tolweb.org/

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