- Arms I 54-65% of ML.
- Arms III 73-103% of ML.
- Arms IV 120-200% of ML.
- Large arm suckers with 10-20 usually separate, pointed to blunt teeth on distal 2/3 of ring.
- Largest suckers not globular.
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- Club length 115 % of ML.
- Club suckers with 11-15 pointed teeth, in contact at bases, over distal 3/4 of ring; enlarged central tooth.
- Sucker stalks divided into two parts with flag-like lateral keel on proximal segment; stalks of suckers in lateral series about twice as long as those of medial series.
- Club protective membranes
- Membranes in two sets with proximal set only slightly broader than distal set.
- Proximal set short(ca. 7% of club length) with 11-13 trabeculae; at broadest point, two trabeculae occur between adjacent lateral suckers
- Distal set long (ca. 93% of club length) with 83-96 broad but separate ( < half the trabecula width), mostly undivided trabeculae (first trabecula may be divided).
Figure. Photomicrograph of the base of the club of C. picteti, Hawaiian waters, preserved. The yellow arrows mark the approximate beginning and end of the proximal protective membrane. The white arrow points to a typical trabecula on the distal protective membrane. Photograph by R. Young.
- Head length 32-50% of ML.
- Head length 32-50% of ML.
- Funnel-locking apparatus: wide, low tragus that slightly undercuts mantle component; broad antitragus that deeply undercuts mantle component.
- Fin length ca. 55% of ML.
- Fin width ca. 45% of ML.
- Eyeball- 3 series of round organs: lateral series= 6 to 9; intermediate series = 8 to 11; medial series = 6 to 10; terminal photophores slightly larger.
- Viscera: two organs.
- Club-tip photophore small (exposed photogenetic surface much less than half of photophore length), with little or no terminal papilla.
- Aboral tentacular stalk with cushion-like photophores.
- Distal-most pad-like photophore partially embedded and located proximal to first club sucker.
- Club-tip photophore with pigment in functional chromatophores.
- All pigment on clubs in chromatophore organs.
- Club sucker stalks without pigment.
- Buccal membrane usually with dense epidermal pigment.
- Olfactory organ with or without chromatophores depending on size.
The very short, proximal segment of the club protective membrane can be difficult to recognize as distinct from the distal segment.
The characters described above are based on several specimens from across the geographical range of C. picteti (off New Zealand, Hawaii, Japan and Indonesia) and the holotype of C. imperator (= synonym) from off Sumatra, Indonesia.
We found considerable geographical variation in the following characters: size of the tentacular club; number of trabeculae proximal to the first club sucker; club sucker dentition; arm-sucker dentition; position of the distal stalk photophores; number of arm suckers; number of ventral arm photophores; sucker size; buccal membrane pigmentation.