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Bicyclus anynana (Butler 1879)

Andrew V. Z. Brower
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Containing group: Bicyclus


Bicyclus anynana has become famous as a model organism for the study of developmental plasticity and molecular genetics of wing pattern formation. It is likely to be the first butterfly species to have its genome sequenced.


Brakefield PM, French V, and Zwaan BJ. 2003. Development and the genetics of evolutionary change within insect species. Annu. Rev. Ecol. Syst. 34: 633-660.

Carroll SB, Gates J, Keys DN, Paddock SW, Panganiban GEF, Selegue JE, and Williams JA. 1994. Pattern formation and eyespot determination in butterfly wings. Science 265: 109-114.

Saccheri IJ, Nichols RA, and Brakefield PM. 2006. Morphological differentiation following experimental bottlenecks in the butterfly Bicyclus anynana (Nymphalidae). Biological Journal- Linnean Society 89: 107-115.

Information on the Internet

Title Illustrations
Click on an image to view larger version & data in a new window
Click on an image to view larger version & data in a new window
Scientific Name Bicyclus anynana
Specimen Condition Live Specimen
Source DOE JGI to sequence genomes of more than 40 new organisms
Copyright © 2005 Antónia Monteiro/University at Buffalo
About This Page

Middle Tennessee State University, Murfreesboro, Tennessee, USA

Correspondence regarding this page should be directed to Andrew V. Z. Brower at

Page: Tree of Life Bicyclus anynana (Butler 1879). Authored by Andrew V. Z. Brower. The TEXT of this page is licensed under the Creative Commons Attribution License - Version 3.0. Note that images and other media featured on this page are each governed by their own license, and they may or may not be available for reuse. Click on an image or a media link to access the media data window, which provides the relevant licensing information. For the general terms and conditions of ToL material reuse and redistribution, please see the Tree of Life Copyright Policies.

Citing this page:

Brower, Andrew V. Z. 2006. Bicyclus anynana (Butler 1879). Version 09 November 2006 (under construction). http://tolweb.org/Bicyclus_anynana/85328/2006.11.09 in The Tree of Life Web Project, http://tolweb.org/

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